Graphium (Pazala) group taxonomy

[cardgane]

Does anyone know of a good recent review of the Pazala (Graphium) group? Looks a capital mess... Chinese literature says one thing, Monastyrskyi states something else... What would be the currently accepted state of taxonomy there? Please, help!

Vadim

[teinopalpus]

www.pemberleybooks.com/product/guide-to-the-butterflies-of-the-palearctic-region-papilionidae-1-tribes-leptocircini-and-teinopalpini/16450/

Try this. I have one.

Jan

[cardgane]

Thank you, Jan!

[Adam Cotton]

Vadim,

I am currently involved in a DNA/morphological study on the subgenus Pazala, which is indeed rather complex. The first paper is at the proof stage so should be published very soon. Of course as soon as it is available I will post news here. The paper will be open access, so everyone will be able to download it free of charge.

The book that Jan recommends is not a bad start, although I say it myself

Adam.

[Adam Cotton]

I thought it would be useful to give a synonymic listing of subgenus Pazala based on the 2009 work:

Genus Graphium Scopoli, 1777 (Type Species: Papilio sarpedon Linnaeus, 1758)
Subgenus Pazala Moore, 1888 (Type Species: Papilio glycerion Gray, 1831)

G. eurous (Leech, 1893)


ssp. caschmirensis (Rothschild, 1895) (Type Locality: North Cashmere)
ssp. sikkimica (Heron, 1899) (Type Locality: Sikkim) ssp. eurous (Leech, 1893) (Type Locality: Chang Yang, Central China)
ssp. panopaea (de Nicéville, 1900) (Type Locality: Tse Kou, Western China)
ssp. melli Racheli & Cotton, 2009 (Type Locality: Yuechengling Shan, Guangxi, China)
ssp. meridionalis (Mell, 1935) (Type Locality: Nordkwangtung: Linping)
ssp. inthanon Katayama, 1986 (Type Locality: Doi Inthanon, Chiang Mai, N. Thailand)
ssp. asakurae (Matsumura, 1907) (Type Locality: Formosa)

G. mandarinus (Oberthür, 1879)


ssp. garhwalica (Katayama, 1988) (Type Locality: Joshimath, 2400 m., Uttar Pradesh, N. W. India)
ssp. paphus (de Nicéville, 1886) (Type Locality: Sikkim)
= glycerion (Gray, 1831) (Nepaul) [Junior Homonym]
ssp. mandarinus (Oberthür, 1879) (Type Locality: Moupin)
ssp. kimurai Murayama, 1982 (Type Locality: Doi Inthanon, N. Thailand)
ssp. fangana (Okano, 1986) (Type Locality: near Fang, Chieng Mai Prov., N. Thailand)

G. tamerlanus (Oberthür, 1876)


ssp. tamerlanus (Oberthür, 1876) (Type Locality: Moupin)
= taliensis (O. Bang-Haas, 1927) (Type Locality: China mer. occ.: Jünnan, Tali)
ssp. kansuensis (O. Bang-Haas, 1933) (Type Locality: Kansu mer. or., Tsinglingschan, Peilingschan)

G. parus (de Nicéville, 1900) (Type Locality: Tse Kou, Western China)


= incertus (O. Bang-Haas, 1927) (Type Locality: China mer. occ.: Tatsienlu)

G. alebion (Gray, [1853]) (Type Locality: Northern China)


= mariesii (Butler, 1881) (Type Locality: Lu-Shan mountains, province of Kiukiang, China)

G. mullah (Alphéraky, 1897)


ssp. mullah (Alphéraky, 1897) (Type Locality: Ja-djóou & Lu-tine, Sé-Tchouen)
= timur (Ney, 1911) (Type Locality: Ta-tsien-lu, S. W. China)
ssp. chungianus (Murayama, 1961) (Type Locality: Gōkan, Zentral-formosa, 2000m.)
= tayal (Shirôzu, 1961) (Type Locality: Vicinity of Urai, N. Formosa)
= chungiyanus (d’Abrera, 1982) [Incorrect Subsequent Spelling]
ssp. kooichii (Morita, 1996) (Type Locality: LacSao, Laos)

G. sichuanica (Koiwaya, 1993) (Type Locality: Dujiang Yan, Sichuan, China)


= hoenei (Mell, 1935) (Type Locality: Chekiang: West- und Ost-Tienmoshan (Fukien, Südkiangsi), Nordkwangtung) [Junior Homonym]

Adam.

[cardgane]

Thank you so much, Adam!!! Just the sort of thing I was looking for (for the start, and then will get the book to study which Jan sent the link to). Without jumping the gun, as it were, may I ask if you expect critical emendations to the 2009 list in your forthcoming paper?

And once again, thank you for the replies and the information!

Vadim

[Adam Cotton]

Jun 19, 2018 2:37:25 GMT -8 cardgane said:

Thank you so much, Adam!!! Just the sort of thing I was looking for (for the start, and then will get the book to study which Jan sent the link to). Without jumping the gun, as it were, may I ask if you expect critical emendations to the 2009 list in your forthcoming paper?

And once again, thank you for the replies and the information!

Vadim

You will have to wait until the paper is published, but I can say that as part of the paper we analysed DNA of the nominate subspecies of all 7 species. Support was excellent (1 for each node) and we found that alebion is basal to the rest, then tamerlanus & parus branched off, with eurous & mullah as a sister pair to mandarinus & sichuanica.

Adam.

[Adam Cotton]

I have added photos of the 7 species above.

Adam.

[Adam Cotton]

As a result of the latest paper detailed here:

insectnet.proboards.com/thread/7638/revision-pazala-mandarinus-group

I can update the classification of subgenus Pazala as follows:

Genus Graphium Scopoli, 1777 (Type Species: Papilio sarpedon Linnaeus, 1758)
Subgenus Pazala Moore, 1888 (Type Species: Papilio glycerion Gray, 1831)

Graphium alebion (Gray, 1853) (Type Locality: Northern China)
= mariesii (Butler, 1881) (Type Locality: Lu-Shan mountains, province of Kiukiang, China)

Graphium tamerlanus (Oberthür, 1876)
ssp. tamerlanus (Oberthür, 1876) (Type Locality: Moupin)
= taliensis (O. Bang-Haas, 1927) (Type Locality: China mer. occ.: Jünnan, Tali)
ssp. kansuensis (O. Bang-Haas, 1933) (Type Locality: Kansu mer. or., Tsinglingschan, Peilingschan)

Graphium parus (de Nicéville, 1900) (Type Locality: Tse Kou, Western China)
= incertus (O. Bang-Haas, 1927) (Type Locality: China mer. occ.: Tatsienlu)

Graphium eurous (Leech, 1893)
ssp. caschmirensis (Rothschild, 1895) (Type Locality: North Cashmere)
= cashmirensis (Moore, 1903) [Incorrect Subsequent Spelling]
= kashmirensis (Bingham, 1907) [Incorrect Subsequent Spelling]
= caschmirense Katayama, 1986 [Incorrect Subsequent Spelling]
ssp. sikkimica (Heron, 1899) (Type Locality: Sikkim)
= sikkima (Moore, 1903) [Incorrect Subsequent Spelling]
= sikhimica (Bingham, 1907) [Incorrect Subsequent Spelling]
ssp. eurous (Leech, 1893) (Type Locality: Chang Yang, Central China)
= eurois (Moore, 1903) [Incorrect Subsequent Spelling]
= euroa (Chou, 1994) [Unjustified Emendation]
ssp. panopaea (de Nicéville, 1900) (Type Locality: Tse Kou, Western China)
ssp. melli Racheli & Cotton, 2009 (Type Locality: Yuechengling Shan, Guangxi, China)
ssp. meridionalis (Mell, 1935) (Type Locality: Nordkwangtung: Linping)
ssp. inthanon Katayama, 1986 (Type Locality: Doi Inthanon, Chiang Mai, N. Thailand)
ssp. asakurae (Matsumura, 1907) (Type Locality: Formosa)
= formosanus (Oberthür, 1909) (Type Locality: Formose: Mont Morrison, 10,000 pieds) [Junior Homonym]
= koxinga (Wileman, 1909) (Type Locality: Formosa) [Junior Homonym]

Graphium mullah (Alphéraky, 1897)
ssp. mullah (Alphéraky, 1897) (Type Locality: Ja-djóou & Lu-tine, Sé-Tchouen)
= timur (Ney, 1911) (Type Locality: Ta-tsien-lu, S. W. China)
= timor (Draeseke, 1923) [Incorrect Subsequent Spelling]
ssp. chungianus (Murayama, 1961) (Type Locality: Gōkan, Zentral-formosa, 2000m.)
= tayal (Shirôzu, 1961) (Type Locality: Vicinity of Urai, N. Formosa)
= chungiyanus (d’Abrera, 1982) [Incorrect Subsequent Spelling]
ssp. kooichii (Morita, 1996) (Type Locality: LacSao, Laos)

Graphium garhwalica (Katayama, 1988) (Type Locality: Joshimath, 2400 m., Uttar Pradesh, N. W. India)

Graphium paphus (de Nicéville, 1886) (Type Locality: Sikkim)
= glycerion (Gray, 1831) (Type Locality: Nepaul) [Junior Homonym]

Graphium sichuanica (Koiwaya, 1993) (Type Locality: Dujiang Yan, Sichuan, China)

Graphium hoeneanus Cotton & Hu, 2018 [Replacement name]
= hoenei (Mell, 1935) (Type Locality: Chekiang: West- und Ost-Tienmoshan (Fukien, Südkiangsi), Nordkwangtung) [Junior Homonym]

Graphium daiyuanae Hu, Zhang & Cotton, 2018 (Type Locality: Sapa (1,600 m), Lao Cai Province, Vietnam)

Graphium confucius Hu, Duan & Cotton, 2018 (Type Locality: Xi Chong (2,000 m), Kunming, C. Yunnan, China)

Graphium mandarinus (Oberthür, 1879)
ssp. stilwelli Cotton & Hu, 2018 (Type Locality: Tacheng (1,900 m), Weixi, W. Yunnan, China)
f. ♂ albarea (Rousseau-Decelle, [1947]) (Type Locality: Ginfu, Etats Schans) [Infrasubspecific]
ssp. mandarinus (Oberthür, 1879) (Type Locality: Moupin)
= mandarina Chou, 1994 [Unjustified Emendation]
ssp. kimurai Murayama, 1982 (Type Locality: Doi Inthanon, N. Thailand)
ssp. fangana (Okano, 1986) (Type Locality: near Fang, Chieng Mai Prov., N. Thailand)

Adam.

[cardgane]

Great work, Adam. Congratulations on the excellent publication and thank you for sharing it with us!