How many subsp does Papilio ulysses has?

[sam9710]

Hi everyone！
This year I decided to collect papilio ulysses subspecies systematically. And I know it have lots of subspecies and may one of a hardest species to make a comprehensive collection. Many subspecies is invalid, so I want to ask for help that how many subsp it has and their locality.

[sam9710]

Here are some of them I’ve got now.

Attachments:

[Adam Cotton]

Here is my list of subspecies of Papilio ulysses [type localities in ( )]:

Papilio ulysses Linnaeus, 1758
ssp. ampelius Rothschild, 1908 (Buru)
ssp. ambelauensis (Schäffler & Schmidbauer 2004) (Ambelau Is.)
ssp. reikoae Kariya, 1988 (Manipa Is.)
ssp. penelope (Dufek, Schäffler & Schmidbauer, 2005) (Kelang Isl.)
ssp. ulysses Linnaeus, 1758 (Asia [= Ambon])
= diomedes Linnaeus, 1758 (Indiis [= Ambon])
f. elisabethae (Dufek, Schäffler & Schmidbauer, 2005) (Pombo Island, east of Ambon)
ssp. scylla (Dufek, Schäffler & Schmidbauer, 2005) (Saparua Isl.)
ssp. dohertius Rothschild, 1898 (Obi)
ssp. telegonus C. Felder & R. Felder, 1860 (Batschian)
= ulyssodes Westwood, 1860 [NN]
= bepe Koçak & Kemal, 2000 [Replacement Name]
ssp. morotaicus Rothschild, 1908 (Mira, Morty I. (= Morotai))
ssp. waterstradti Goode, 2012 (Moluques, Ile Gebi [= Gebe Island])
ssp. dirce Jordan, 1909 (Misol)
ssp. physkon Fruhstorfer, 1909 (Waigiu)
ssp. calypso (Schäffler & Schmidbauer 2004) (Salawati)
ssp. denticulatus Joicey & Talbot, 1916 (Biak, Schouten Is.)
ssp. autolycus C. Felder & R. Felder, 1865 (Nova Guinea: Dorey)
= penelope Wallace, 1865 (New Guinea, Waigiou, Aru Is.) [JH]
= eugenius Fruhstorfer, 1909 (Deutsch-New Guinea)
= taxiles Fruhstorfer, 1909 (British Neu-Guinea, Milnebai)
= altovolensis (Schäffler & Schmidbauer, 2004) (Arfak Mts., Irian Jaya)
aberr. ♀ conjuncta Hagen, 1897 (Neu-Guinea)
aberr. ♀ transiens Hagen, 1897 (Neu-Guinea)
f. ♀ principalis Fruhstorfer, 1909 (German New Guinea)
f. ♀ roxane Fruhstorfer, 1909 (Deutsch-Neu-Guinea)
= roxana Jordan, 1909 [ISS]
f. ♀ senidae (Schäffler & Schmidbauer 2004) (Arfak Mts., Irian Jaya)
ssp. oxyartes Fruhstorfer, 1909 (Aru Island)
ssp. melanotica Hagen, 1897 (Dampier)
= karkarensis (Schäffler & Schmidbauer 2004) (Karkar Is., Papua New Guinea)
ssp. telemachus Montrouzier, 1856 (Woodlark)
= telephanes Fruhstorfer, 1902 (Kiriwina and Fergusson)
= agasophus Fruhstorfer, 1903 (Fergusson, D’Entrecasteaux-Inseln)
ssp. joesa Butler, 1869 (Queensland)
ssp. gabrielis Rothschild, 1898 (St. Gabriel, Admiralty Islands)
ssp. kallinikos Fruhstorfer, 1903 (Neu-Mecklenburg = Neu-Irland)
= jennifeae Jakusch, 2007 (Schleinitz Mts., Labusar, 200-550 m, New Ireland)
ssp. ambiguus Rothschild, 1895 (New Britain)
= marius Fruhstorfer, 1903 (Mioko, Neu-Lauenburg)
ssp. nigerrimus Ribbe, 1898 (Schortlands, deutschen Salomo-Inseln)
f. ♀ nigerrimus Ribbe, 1898 (Schortlands, deutschen Salomo-Inseln)
f. ♀ cyanippe Jordan, 1909 (Bougainville and Shortland Islands)
ssp. rothschildianus Fruhstorfer, 1909 [Replacement Name]
= georgius Rothschild, 1908 (Solomon Islands: Guizo) [JH]
ssp. orsippus Godman & Salvin, 1888 (Solomon Islands, Aola in Guadalcanar Island)
f. ♀ hilaria Jordan, 1909 (Guadalcanal)
f. ♀ penthina Jordan, 1909 (Isabel)

ISS = Incorrect Subsequent Spelling
JH = Junior Homonym
NN = Nomen Nudum

[Adam Cotton]

Note that some names treated as subspecies above may actually be synonyms. If anyone notices any errors please post them here for our information.

Adam.

[exoticimports]

ssp. nigerrimus Ribbe, 1898 (Schortlands, deutschen Salomo-Inseln)

Tennent does not recognize that as a valid ssp in fact states that though there are regional (island) forms, there exist sufficient intermediary grades to determine all one ssp.

Chuck

[Adam Cotton]

Feb 5, 2019 4:57:33 GMT -8 exoticimports said:

ssp. nigerrimus Ribbe, 1898 (Schortlands, deutschen Salomo-Inseln)

Tennent does not recognize that as a valid ssp in fact states that though there are regional (island) forms, there exist sufficient intermediary grades to determine all one ssp.

Chuck

Thank you for pointing that out.

Tennent (2002 - Butterflies of the Solomon Islands) does indeed treat nigerrimus Ribbe, 1898 as a synonym of orsippus Godman & Salvin, 1888.

Adam.

[jhyatt]

I have a hard-and-fast rule about recognizing doubtful subspecies in my collection: If I lack an example, I consider it to be a synonym anyway. But If I have a specimen, then
I say it's valid and label it as such!

Heh heh,

jh

[exoticimports]

Feb 5, 2019 5:42:43 GMT -8 jhyatt said:

I have a hard-and-fast rule about recognizing doubtful subspecies in my collection: If I lack an example, I consider it to be a synonym anyway. But If I have a specimen, then
I say it's valid and label it as such!

Heh heh,

jh

Call me awful, but I have a rule too: don't bother recording the name for anything anymore, it changes constantly. Date & location don't change, at least that often.

Chuck

[jhyatt]

I put most everything into unit trays. I use a tray label for the species or ssp, but I seldom put individual determination labels on specimens.
But it's a rare thing for me to update a label with the most current name... I can remember the old ones, but not the new! I haven't changed country names on
locality labels, either... I have obsolete places like USSR and Northwest Territories and certain English counties on my labels, and am happy to leave them so.

Cheers,
jh

[coloradeo]

Do you have any species labeled “The New World”?

[jhyatt]

No, but more than once I'd had an old world specimen turn up in a batch of papered neotropical bugs.. Never understood how that can happen...
jh

[sam9710]

Feb 5, 2019 4:01:51 GMT -8 Adam Cotton said:

Here is my list of subspecies of Papilio ulysses [type localities in ( )]:

Papilio ulysses Linnaeus, 1758
ssp. ampelius Rothschild, 1908 (Buru)
ssp. ambelauensis (Schäffler & Schmidbauer 2004) (Ambelau Is.)
ssp. reikoae Kariya, 1988 (Manipa Is.)
ssp. penelope (Dufek, Schäffler & Schmidbauer, 2005) (Kelang Isl.)
ssp. ulysses Linnaeus, 1758 (Asia [= Ambon])
= diomedes Linnaeus, 1758 (Indiis [= Ambon])
f. elisabethae (Dufek, Schäffler & Schmidbauer, 2005) (Pombo Island, east of Ambon)
ssp. scylla (Dufek, Schäffler & Schmidbauer, 2005) (Saparua Isl.)
ssp. dohertius Rothschild, 1898 (Obi)
ssp. telegonus C. Felder & R. Felder, 1860 (Batschian)
= ulyssodes Westwood, 1860 [NN]
= bepe Koçak & Kemal, 2000 [Replacement Name]
ssp. morotaicus Rothschild, 1908 (Mira, Morty I. (= Morotai))
ssp. waterstradti Goode, 2012 (Moluques, Ile Gebi [= Gebe Island])
ssp. dirce Jordan, 1909 (Misol)
ssp. physkon Fruhstorfer, 1909 (Waigiu)
ssp. calypso (Schäffler & Schmidbauer 2004) (Salawati)
ssp. denticulatus Joicey & Talbot, 1916 (Biak, Schouten Is.)
ssp. autolycus C. Felder & R. Felder, 1865 (Nova Guinea: Dorey)
= penelope Wallace, 1865 (New Guinea, Waigiou, Aru Is.) [JH]
= eugenius Fruhstorfer, 1909 (Deutsch-New Guinea)
= taxiles Fruhstorfer, 1909 (British Neu-Guinea, Milnebai)
= altovolensis (Schäffler & Schmidbauer, 2004) (Arfak Mts., Irian Jaya)
aberr. ♀ conjuncta Hagen, 1897 (Neu-Guinea)
aberr. ♀ transiens Hagen, 1897 (Neu-Guinea)
f. ♀ principalis Fruhstorfer, 1909 (German New Guinea)
f. ♀ roxane Fruhstorfer, 1909 (Deutsch-Neu-Guinea)
= roxana Jordan, 1909 [ISS]
f. ♀ senidae (Schäffler & Schmidbauer 2004) (Arfak Mts., Irian Jaya)
ssp. oxyartes Fruhstorfer, 1909 (Aru Island)
ssp. melanotica Hagen, 1897 (Dampier)
= karkarensis (Schäffler & Schmidbauer 2004) (Karkar Is., Papua New Guinea)
ssp. telemachus Montrouzier, 1856 (Woodlark)
= telephanes Fruhstorfer, 1902 (Kiriwina and Fergusson)
= agasophus Fruhstorfer, 1903 (Fergusson, D’Entrecasteaux-Inseln)
ssp. joesa Butler, 1869 (Queensland)
ssp. gabrielis Rothschild, 1898 (St. Gabriel, Admiralty Islands)
ssp. kallinikos Fruhstorfer, 1903 (Neu-Mecklenburg = Neu-Irland)
= jennifeae Jakusch, 2007 (Schleinitz Mts., Labusar, 200-550 m, New Ireland)
ssp. ambiguus Rothschild, 1895 (New Britain)
= marius Fruhstorfer, 1903 (Mioko, Neu-Lauenburg)
ssp. nigerrimus Ribbe, 1898 (Schortlands, deutschen Salomo-Inseln)
f. ♀ nigerrimus Ribbe, 1898 (Schortlands, deutschen Salomo-Inseln)
f. ♀ cyanippe Jordan, 1909 (Bougainville and Shortland Islands)
ssp. rothschildianus Fruhstorfer, 1909 [Replacement Name]
= georgius Rothschild, 1908 (Solomon Islands: Guizo) [JH]
ssp. orsippus Godman & Salvin, 1888 (Solomon Islands, Aola in Guadalcanar Island)
f. ♀ hilaria Jordan, 1909 (Guadalcanal)
f. ♀ penthina Jordan, 1909 (Isabel)

ISS = Incorrect Subsequent Spelling
JH = Junior Homonym
NN = Nomen Nudum

Thank you Adam！It did a great help.

[sam9710]

 I still have several specimens need identify，if someone can help me.
This one if from ：Timoenia Mauns Is Manus prov PNG

[sam9710]

 here is the underside. I can not make sure his ssp.

[sam9710]

 female from Bulolo（Buioio？）Morobe prov PNG. I have ssp.gabrielis male with the same collecting data，is this one a gabrielis female？