Revision of G. (Pazala) mandarinus group

[Adam Cotton]

I am happy to announce a new paper was just published today:

Revision of Pazala Moore, 1888: The Graphium (Pazala) mandarinus (Oberthür, 1879) Group, with Treatments of Known Taxa and Descriptions of New Species and New Subspecies (Lepidoptera: Papilionidae)

Authors:

SHAO-JI HU, ADAM M. COTTON, FABIEN L. CONDAMINE, KUANG DUAN, RONG-JIANG WANG, YU-FENG HSU, XIN ZHANG, JUN CAO

Abstract:

The previously recognised closely related species Graphium (Pazala) mandarinus (Oberthür, 1879) and G. (P.) sichuanica (Koiwaya, 1993) are shown to comprise seven species as a result of both molecular and morphological analysis. Molecular dating analysis is also performed on the mandarinus group in order to investigate the divergence time of the taxa. Two taxa, G. (P.) garhwalica (Katayama, 1988) stat. nov. and G. (P.) paphus (de Nicéville, 1886) stat. nov., are raised from subspecific to specific status; G. (P.) hoeneanus Cotton & Hu nom. nov., stat. rev. is separated from sichuanica at species level; and two previously unrecognised new species, G. (P.) daiyuanae Hu, Zhang & Cotton sp. nov. and G. (P.) confucius Hu, Duan & Cotton sp. nov. are described from Vietnam and China respectively, the latter being sympatric with nominate G. (P.) mandarinus. The identity of the lectotype of G. (P.) mandarinus is confirmed and a lectotype is designated for the taxon Papilio Glycerion Gray, 1831. A new subspecies of G. (P.) mandarinus is described from western Yunnan and northern Myanmar, G. (P.) mandarinus stilwelli Cotton & Hu ssp. nov.

The paper is open access and can be downloaded here:

www.mapress.com/j/zt/article/view/zootaxa.4441.3.1/15223

Note the file is 24MB in size.

Adam.

[Adam Cotton]

Since the paper is open access I can post photos of specimens from the figures here.

Firstly Graphium garhwalica and paphus were found to be separate species to each other and to mandarinus. Here's the plate showing specimens and how to distinguish them:


G. (P.) garhwalica (Katayama, 1988) stat. nov. (A–B) and G. (P.) paphus (de Nicéville, 1886) stat. nov. (C–F); upperside in the first row, underside in the second row; scale bar = 10 mm. A: ♂, Joshimath, N.W. India, PARATYPE; B: ♀, ditto; C: ♂, Tiger Hill, Darjeeling, N. India; D: ♀, ditto; E: ♂, Metok, S. Tibet, China; F: ♂, Tarung Hka River, N. Sagaing, Myanmar.

Also Graphium sichuanica from Sichuan was shown to be separate from the taxon previously known as hoenei, which is a junior homonym and was renamed hoeneanus, which is found in SE China.


G. (P.) sichuanica (Koiwaya, 1993) (A–C) and G. (P.) hoeneanus Cotton & Hu nom. nov., stat. rev. (D–F); upperside in the first row, underside in the second row; scale bar = 10 mm. A and B: ♂, Laohegou, Pingwu, Sichuan, W. China, © Peking University & Laohegou Nature Reserve; C: ♀, Tangjiahe, Qingchuan, Sichuan, W. China, © Peking University & Tangjiahe Nature Reserve; D: ♂, W. Tianmu Shan (350 m), Zhejiang, E. China; E: ♀, Tiantong (400 m), Ningbo, Zhejiang, E. China, © Jian-Qing Zhu (Shanghai Zoological Park, Shanghai, China); F: ♂, Lingui (360 m), Guilin, Guangxi, S.W. China.


Two new species were named:


Graphium (Pazala) daiyuanae Hu, Zhang & Cotton, sp. nov. holotype, ♂ Sapa (1,600 m), Lao Cai Province, N. Vietnam, 2015–VI


Graphium (Pazala) confucius Hu, Duan & Cotton sp. nov. holotype, ♂, Xi Chong (2,000 m), Kunming, C. Yunnan, China, 2015–V–31

One new subspecies was also named:


Graphium (Pazala) mandarinus stilwelli Cotton & Hu ssp. nov. holotype, ♂, Tacheng (1,900 m), Weixi, W. Yunnan, China, 2015–IV–29

Adam.

[exoticimports]

I wonder how much total time was put into this wonderful paper.

[LEPMAN]

Its a very nice paper indeed!

[Adam Cotton]

Jun 29, 2018 16:51:02 GMT -8 exoticimports said:

I wonder how much total time was put into this wonderful paper.

3 years.